Order Suborder Family Family Distance BIN Distance BIN n

Order Suborder Family Family Distance BIN Distance BIN n

S3 Table. Mean Dayhoff distance (+/- SE) based on amino acid divergences in the barcode
region of COI for ten orders of arachnids with data for more than one family. Values are
reported for the divergences among families in each order (Family Distance) and among the BINs in
each family (BIN Distance). All Distances > 0.25 are shown in bold, while families highlighted in
green are those which included at least one species with an indel.
Order
Suborder
Solifugae
Family
Family Distance
0.071 (SE = 0.001)
Rhagodidae
0.069
Galeodidae
0.070
Ammotrechidae
0.072
Eremobatidae
0.074
Amblypygi
0.084 (SE = 0.003)
Euamblypygi
Phrynichidae
0.076
Euamblypygi
Charontidae
0.081
Euamblypygi
Phrynidae
0.086
Euamblypygi
Charinidae
0.092
Scorpiones
0.099 (SE = 0.010)
Euscorpiidae
0.080
Vaejovidae
0.078
Chactidae
0.086
Scorpionidae
0.094
Hemiscorpiidae
0.114
Buthidae
0.144
Araneae
0.158 (SE = 0.003)
Araneomorphae Corinnidae
0.123
Araneomorphae Desidae
0.124
Araneomorphae Nephilidae
0.125
Araneomorphae Nesticidae
0.132
Araneomorphae Stiphidiidae
0.130
Araneomorphae Dictynidae
0.132
Araneomorphae Pimoidae
0.135
Araneomorphae Ctenidae
0.136
Araneomorphae Gnaphosidae
0.135
Araneomorphae Sparassidae
0.136
Araneomorphae Araneidae
0.136
Araneomorphae Liocranidae
0.135
Araneomorphae Mimetidae
0.137
Araneomorphae Mysmenidae
0.136
Araneomorphae Agelenidae
0.140
Araneomorphae Hahniidae
0.138
Araneomorphae Linyphiidae
0.139
Araneomorphae Mecysmaucheniidae 0.138
Araneomorphae Thomisidae
0.139
Araneomorphae Trachelidae
0.140
BIN Distance
0.023 (SE = NA)
0.023
NA
NA
NA
0.051 (SE = 0.007)
0.038
0.053
0.062
NA
0.036 (SE = 0.012)
0.000
0.015
0.062
0.051
NA
0.053
0.064 (SE = 0.005)
0.007
0.059
0.039
0.048
NA
0.062
0.024
NA
0.068
0.057
0.035
0.012
0.053
0.057
0.079
0.086
0.050
0.064
0.032
0.087
BIN n
2
1
1
1
2
3
2
1
2
52
3
22
1
47
3
3
13
10
1
42
4
1
49
14
97
3
4
14
31
16
348
5
59
3
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Mygalomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Mygalomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Araneomorphae
Mygalomorphae
Araneomorphae
Mygalomorphae
Araneomorphae
Mygalomorphae
Araneomorphae
Araneomorphae
Mygalomorphae
Araneomorphae
Araneomorphae
Mygalomorphae
Mygalomorphae
Araneomorphae
Cybaeidae
Miturgidae
Theridiidae
Anyphaenidae
Pisauridae
Theridiosomatidae
Uloboridae
Lycosidae
Salticidae
Stenochilidae
Archaeidae
Eutichuridae
Eresidae
Philodromidae
Tetragnathidae
Phrurolithidae
Psechridae
Zoropsidae
Segestriidae
Clubionidae
Dysderidae
Nemesiidae
Amaurobiidae
Periegopidae
Phyxelididae
Caponiidae
Dipluridae
Diguetidae
Filistatidae
Palpimanidae
Hypochilidae
Ctenizidae
Huttoniidae
Actinopodidae
Anapidae
Antrodiaetidae
Pholcidae
Symphytognathidae
Migidae
Leptonetidae
Scytodidae
Theraphosidae
Mecicobothriidae
Telemidae
0.139
0.137
0.140
0.142
0.144
0.144
0.147
0.146
0.144
0.145
0.148
0.148
0.149
0.150
0.150
0.148
0.151
0.162
0.158
0.156
0.160
0.158
0.159
0.160
0.161
0.164
0.166
0.169
0.172
0.175
0.176
0.182
0.182
0.187
0.194
0.193
0.191
0.187
0.193
0.196
0.198
0.198
0.197
0.205
0.063
NA
0.080
0.047
0.032
0.103
0.066
0.045
0.057
NA
0.079
0.078
NA
0.061
0.064
0.023
0.082
NA
0.058
0.039
0.062
0.048
0.090
NA
0.088
NA
0.112
NA
NA
NA
NA
0.058
NA
0.047
0.181
NA
0.123
0.155
0.083
0.059
NA
0.082
NA
NA
24
1
93
7
12
2
5
101
89
1
14
5
1
31
71
7
44
1
2
38
34
13
13
1
25
1
2
1
1
1
1
2
1
18
3
1
84
5
27
2
1
5
1
1
Araneomorphae Sicariidae
Mesothelae
Liphistiidae
Ixodida
Ixodoidea
Ixodoidea
Ixodoidea
Sarcoptiformes
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Argasidae
Ixodidae
Nuttalliellidae
Cepheidae
Chamobatidae
Eremaeidae
Mochlozetidae
Tenuialidae
Tegoribatidae
Phenopelopidae
Ceratozetidae
Mycobatidae
Achipteriidae
Peloppiidae
Megeremaeidae
Hydrozetidae
Scutoverticidae
Cymbaeremaeidae
Oppiidae
Eremulidae
Haplozetidae
Oribatulidae
Malaconothridae
Eremobelbidae
Astegistidae
Oribatellidae
Trhypochthoniidae
Hermanniidae
Ameronothridae
Gymnodamaeidae
Gustaviidae
Suctobelbidae
Neoliodidae
Galumnidae
Quadroppiidae
Crotoniidae
Hermanniellidae
Euzetidae
Parakalummidae
Liacaridae
0.227
0.246
0.213 (SE = 0.008)
0.206
0.203
0.229
0.250 (SE = 0.010)
0.173
0.176
0.176
0.176
0.177
0.178
0.178
0.178
0.179
0.179
0.179
0.180
0.180
0.180
0.181
0.181
0.183
0.183
0.184
0.184
0.186
0.186
0.188
0.189
0.190
0.192
0.193
0.194
0.194
0.194
0.195
0.198
0.200
0.200
0.202
0.203
0.204
0.006
NA
0.097 (SE = 0.017)
0.114
0.080
NA
0.046 (SE = 0.005)
0.008
0.000
0.020
0.014
NA
0.032
0.021
0.026
0.037
0.037
0.020
NA
0.023
0.028
0.044
0.024
NA
0.045
0.024
0.006
NA
NA
0.065
0.011
0.004
0.034
0.013
0.064
0.021
0.028
0.089
0.010
0.048
NA
0.005
0.058
0.101
3
1
4
67
1
8
3
28
7
1
3
12
104
11
9
18
1
2
39
15
39
1
7
40
8
1
1
5
10
2
3
9
2
12
3
20
4
35
1
2
5
5
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Endeostigmata
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Oribatida
Endeostigmata
Astigmata
Endeostigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Astigmata
Oribatida
Astigmata
Astigmata
Astigmata
Endeostigmata
Astigmata
Mesostigmata
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Sejida
Nothridae
Scheloribatidae
Damaeidae
Tectocepheidae
Thyrisomidae
Oripodidae
Passalozetidae
Alicorhagiidae
Oribotritiidae
Eniochthoniidae
Brachychthoniidae
Euphthiracaridae
Carabodidae
Hypochthoniidae
Alycidae
Alloptidae
Nanorchestidae
Acaridae
Pterolichidae
Proctophyllodidae
Pyroglyphidae
Avenzoariidae
Xolalgidae
Trouessartiidae
Psoroptidae
Carpoglyphidae
Gabuciniidae
Analgidae
Pteronyssidae
Histiostomatidae
Listrophoridae
Phthiracaridae
Syringobiidae
Myocoptidae
Knemidokoptidae
Terpnacaridae
Sarcoptidae
Paratennulidae
Parasitidae
Arctacaridae
Microgyniidae
Oplitidae
Sejidae
0.206
0.208
0.212
0.212
0.227
0.232
0.239
0.240
0.241
0.242
0.254
0.273
0.278
0.282
0.310
0.316
0.321
0.322
0.322
0.324
0.326
0.327
0.331
0.332
0.332
0.334
0.335
0.337
0.338
0.341
0.348
0.350
0.377
0.378
0.385
0.406
0.460
0.229 (SE = 0.009)
0.179
0.182
0.187
0.188
0.188
0.188
0.012
0.071
0.067
0.076
0.141
0.038
NA
0.004
NA
0.004
0.051
0.149
0.140
NA
0.119
0.025
0.133
0.034
0.016
0.018
0.022
0.072
0.040
0.027
0.039
NA
0.016
0.037
0.052
0.087
0.093
0.174
NA
NA
0.044
0.061
NA
0.120 (SE = 0.017)
NA
0.059
0.080
0.057
NA
0.036
13
34
16
51
3
6
1
5
1
2
66
4
2
1
5
2
9
4
1
52
7
7
3
13
2
1
2
5
7
2
3
9
1
1
2
3
1
1
54
2
4
1
2
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Monogynaspida
Pseudoscorpiones
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Iocheirata
Epiocheirata
Epiocheirata
Epiocheirata
Iocheirata
Iocheirata
Epiocheirata
Epiocheirata
Iocheirata
Trombidiformes
Prostigmata
Prostigmata
Prostigmata
Urodinychidae
Trematuridae
Ologamasidae
Pachylaelapidae
Dinychidae
Trachytidae
Macrochelidae
Digamasellidae
Melicharidae
Veigaiidae
Varroidae
Ascidae
Zerconidae
Laelapidae
Phytoseiidae
Blattisociidae
Ameroseiidae
Dermanyssidae
Hyidae
Geogarypidae
Larcidae
Cheliferidae
Olipiidae
Ideoroncidae
Syarinidae
Gymnobisiidae
Neobisiidae
Atemnidae
Garypidae
Parahyidae
Chernetidae
Chthoniidae
Feaellidae
Tridenchthoniidae
Sternophoridae
Withiidae
Lechytiidae
Pseudogarypidae
Cheiridiidae
Protziidae
Hydryphantidae
Sperchontidae
0.195
0.199
0.199
0.204
0.215
0.218
0.223
0.231
0.231
0.236
0.247
0.246
0.255
0.289
0.285
0.283
0.315
0.316
0.331 (SE = 0.004)
0.304
0.307
0.305
0.311
0.311
0.311
0.318
0.323
0.322
0.327
0.328
0.346
0.340
0.338
0.349
0.340
0.341
0.338
0.363
0.365
0.370
0.341 (SE = 0.013)
0.238
0.251
0.250
0.035
0.035
0.073
0.178
0.084
NA
0.119
0.198
0.106
0.093
NA
0.157
0.183
0.273
0.234
0.201
0.078
NA
0.190 (SE = 0.017)
NA
0.083
NA
0.141
0.214
NA
0.263
NA
0.172
0.157
0.164
NA
0.172
0.194
NA
NA
0.171
NA
NA
0.251
0.298
0.111 (SE = 0.012)
NA
0.025
0.035
5
12
12
3
9
1
14
47
20
3
1
51
14
30
94
26
7
1
1
2
1
6
11
1
5
1
8
6
2
1
14
20
1
1
2
1
1
2
3
1
8
5
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Prostigmata
Lebertiidae
Torrenticolidae
Thyasidae
Mideopsidae
Pionidae
Acalyptonotidae
Arrenuridae
Unionicolidae
Eylaidae
Anystidae
Rhagidiidae
Limnesiidae
Bdellidae
Hydrachnidae
Ereynetidae
Eupodidae
Trombidiidae
Penthaleidae
Penthalodidae
Tydeidae
Trombiculidae
Siteroptidae
Scutacaridae
Erythraeidae
Hygrobatidae
Calyptostomatidae
Cunaxidae
Stigmaeidae
Pyemotidae
Microtrombidiidae
Tarsonemidae
Cheyletidae
Eriophyidae
Tetranychidae
Tenuipalpidae
Dyspnoi
Euponoi
Laniatores
Laniatores
Laniatores
Laniatores
Cyphophthalmi
Dyspnoi
Ceratolasmatidae
Phalangiidae
Gonyleptidae
Stygnopsidae
Stygnommatidae
Epedanidae
Troglosironidae
Trogulidae
Opiliones
0.255
0.252
0.256
0.264
0.263
0.273
0.275
0.279
0.298
0.291
0.290
0.296
0.304
0.312
0.323
0.337
0.335
0.342
0.342
0.344
0.345
0.351
0.350
0.374
0.388
0.390
0.408
0.399
0.397
0.417
0.449
0.459
0.479
0.522
0.574
0.354 (SE = 0.010)
0.308
0.308
0.316
0.316
0.321
0.322
0.324
0.329
0.017
0.020
0.055
0.034
0.046
NA
0.042
0.059
0.016
0.085
0.042
0.075
0.125
0.042
0.108
0.120
0.098
0.097
0.104
0.192
0.180
0.096
0.098
0.200
0.292
0.086
0.181
0.313
0.167
0.181
0.171
0.117
0.128
0.168
0.176
0.108 (SE = 0.029)
0.013
0.027
NA
0.155
0.037
0.004
0.025
NA
7
7
4
3
16
1
16
13
6
61
72
10
65
2
5
177
12
7
12
62
22
30
24
126
10
2
22
31
2
14
17
4
9
39
13
8
4
1
3
4
2
14
1
Laniatores
Laniatores
Euponoi
Cyphophthalmi
Laniatores
Cyphophthalmi
Laniatores
Laniatores
Cyphophthalmi
Laniatores
Laniatores
Cyphophthalmi
Kimulidae
Biantidae
Sclerosomatidae
Stylocellidae
Zalmoxidae
Pettalidae
Samoidae
Icaleptidae
Neogoveidae
Triaenonychidae
Podoctidae
Sironidae
0.331
0.331
0.333
0.345
0.357
0.359
0.380
0.390
0.404
0.420
0.421
0.492
NA
NA
0.051
0.105
NA
0.120
NA
NA
0.266
0.195
NA
0.300
1
1
7
10
1
67
1
1
24
4
1
20