THE TEN THOUSAND ISLANDS COAST OF FLORIDA: A MODERN
Transcrição
THE TEN THOUSAND ISLANDS COAST OF FLORIDA: A MODERN
Proceedings of the 5th International Conference on Coastal Sediments, 5:1773-1784, R. A., Davis, Jr., Ed., May, 2003, Clearwater Beach, Florida © 2003 by the 5th International Conference on Coastal Sediments THE TEN THOUSAND ISLANDS COAST OF FLORIDA: A MODERN ANALOG TO LOW-ENERGY MANGROVE COASTS OF CRETACEOUS EPEIRIC SEAS Kenneth J. Lacovara,1 Jennifer R. Smith,2 Joshua B. Smith,3 and Matthew C. Lamanna4 Abstract: Epeiric seas stretched across vast regions of continents during the eustatic highs of the Late Cretaceous. Along these seas, low-energy conditions prevailed. Halophytic plants, including the tree-fern Weichselia reticulata, colonized these quiescent coasts, forming highly productive mangrove ecosystems. The morphodynamic behavior of Cretaceous epeiric coasts can be better understood though analogy with similar environments that exist today. The Ten Thousand Islands coast of southwestern Florida provides an ideal analog. The low-energy wave regime and attenuated tidal range is probably representative of conditions that prevailed along Cretaceous epeiric coasts. The invasion of the littoral zone along the Ten Thousand Islands coast by robust mangrove vegetation has led to a bimodal response to Holocene sea level rise. Both transgressive and regressive behaviors occur simultaneously along different reaches, creating a very complex stratigraphic record. Late Cretaceous coastal deposits exposed in the Bahariya Oasis of Egypt record the development of a similar low-energy mangrove coast in response to rising Cenomanian sea level. These deposits record coeval transgressive and regressive sequences that appear to have resulted from: 1) heterogeneities in the paleolittoral system, and; 2) colonization of the littoral zone by mangrove vegetation, that includes Weichselia reticulata and possibly other mangrove species. Study of the modern analog provided by the Ten Thousand Islands coast has facilitated a more complete understanding of the paleomorphodynamics of low-energy Mesozoic epeiric coasts. Conversely, the study of ancient coastal deposits, laid down in Cretaceous “hothouse” conditions, holds implications for understanding coastal response to modern global warming and sea level rise. INTRODUCTION The sea level record for most of Earth history is completely unknown. Meaningful estimates extend back only to about the Paleozoic/Mesozoic boundary (245 Ma) (Haq et al., 1987) and represent only the last 5% of Earth time. Sea level is currently anomalously low and has been much higher during most of the post-Paleozoic. Peak levels occurred during the Late Cretaceous (99-65 Ma) and may have reached 300 m above present. The eustatic highs of the Late Cretaceous caused marine inundation of vast terrestrial areas, forming epeiric seas over part of every continent (Figure 1). Epeiric seas generally took the form of lobate coastal bights or elongate seaways. Coasts along these seas would have been unlike most of today’s open-ocean coasts. Fetch along these constricted bodies of water would have been quite limited in most directions. The narrow geometry of epeiric seas would have attenuated tidal flow in 1 Drexel University, 251 Curtis Hall, 3141 Chestnut Street, Philadelphia, PA 19104; [email protected] 2 Washington University, Campus Box 1169, 1 Brookings Drive, St. Louis, MO 63130; [email protected] 3 Washington University, Campus Box 1169, 1 Brookings Drive, St. Louis, MO 63130; [email protected] 4 University of Pennsylvania, 240 South 33rd St., Philadelphia, PA 19104; [email protected] 1773 Lacovara et al., 2003 many cases. The resulting energy state of most epeiric coasts would have been much lower than the average state along coasts today. Figure 1. Cenomanian paleogeographic reconstruction for 94 Ma. The paleolocation of Bahariya Formation deposits is shown in red. These deposits were situated along the Bahariya Bight, a sheltered embayment off the Tethys Sea. Figure modified from Scotese (2001). Gently sloping coastal plain regions were most likely to be inundated by rising Cretaceous seas. Because the width of back-barrier paralic wetlands is inversely related to the gradient of the surface being transgressed, immense swaths of coastal wetlands were produced. Broad tidal flats, in lowenergy conditions, in the equable “hothouse” Cretaceous climate provided ideal conditions for phytoproductivity in coastal wetlands. Plants in epeiric coastal biomes benefited from frost-free conditions extending into high latitudes and played an important role in determining the response of Cretaceous coasts to fluctuations in sea level. To understand the coastal morphodynamics of ancient inland seas, appropriate modern analogs must be examined. An ideal proxy would be a very low-energy marine coast, with high plant productivity, along a very gentle continental shelf/coastal plain gradient. The Ten Thousand Islands coast of southwestern Florida provides an ideal example. 1774 Lacovara et al., 2003 THE TEN THOUSAND ISLANDS The shelf off the southern Florida coast is approximately 200 km wide, with shoreface gradients less than 1:3,500. These conditions, along with limited fetch across the Gulf of Mexico, result in a mean annual wave height of only 10 to 25 cm (Tanner, 1960). Although tides along this coast are microtidal, with spring tides less than 1.3 m, they dominate the weak wave regime (Davis et al., 1992). Placid conditions along this reach have promulgated the development of a mangrove coast between the cuspate forelands of Cape Romano and Cape Sable. The Ten Thousand Islands lie along the northern portion of the mangrove coast and consist of a great number of mangals (mangrove vegetated tidal flats) separated by sandy tidal channels (Figure 2). Figure 2. The Ten Thousand Islands lie along the northern part of the mangrove coast of southwestern Florida. (NASA satellite image) A variety of mangrove species and mangrove-associates colonize the Ten Thousand Islands. Dominant among them is the angiosperm Rhizophora mangle (red mangrove) (Figure 3). Rhizophora is an obligate of the paralic realm and specifically adapted to its stresses. For example, its many stilt roots buttress it from wave attack. It obtains fresh water by filtering salt water though its roots, a process that produces xylem sap that is 100 times less saline than sea water (Thomlinson, 1986). To conserve “manufactured” water, the leaves of Rhizophora possess many xeromorphic features usually associated with desert vegetation. Most significantly, with respect to geomorphology, Rhizophora is capable of colonizing the littoral zone by the lateral extension of its root system and through the production of floating, viviparous seedlings. 1775 Lacovara et al., 2003 Figure 3. Rhizophora mangle (red mangrove) growing on the shoreface along the Ten Thousand Islands coast. (Photo: K. Lacovara) Negligible wave-energy and the growth of sturdy mangrove vegetation on the foreshore have created an interesting mode of coastal evolution along parts of Florida’s mangrove coast – seaward progradation of the coast during sea level rise. Enos and Perkins (1979) found that the mangrove coast at Cape Sable prograded up to 8 km seaward, during the later portion of the Holocene. They concluded that the coast in this area switched from transgressive to regressive mode following a deceleration in sea level rise. Parkinson (1987; 1989) found similar evidence from a series of vibracores taken in the Ten Thousand Islands. He generalized the stratigraphy into three sequences: transgressive, transitional, and regressive. He also attributed the change in modality to decelerating Holocene sea level rise, which he concluded, based on Scholl (1969), occurred between 3,500 to 3,200 BP. The regressive sequence Parkinson (1989) presented (Figure 4) is interesting in that it showed marine shoreface deposits under mangrove peats. This relationship does not conform to classic regressive or transgressive models (see respectively: Bernard and LeBlanc, 1965, and; Kraft, 1971) in which paralic deposits are shown separated from marine deposits by a barrier facies in conformable sequences. Parkinson’s sequence seems at first a violation of Walther’s Law. However, the classic models are based on the response of sandy coasts to moderate- to high-energy conditions and appear not to apply to very low-energy vegetated coasts. The coastal setting represented by the Ten Thousand Islands is, today, rare. However, along the warm, low-energy coasts of Cretaceous epeiric seas, analogous physical settings must have been common. The model presented by Parkinson (1989), therefore, is important to consider when examining Mesozoic coastal sediments. 1776 Lacovara et al., 2003 Figure 4. Low-energy mangrove coast. This unusual stratigraphy is formed as mangrove vegetation colonizes the shoreface. The coast migrates seaward and upward, even as sea level rises. Note that the vertical sequence (1-4) does not follow the horizontal sequence (1-4), an apparent violation of Walther’s Law. (Modified and idealized after Parkinson, 1989). THE BAHARIYA BIGHT During the sea level highs of the Late Cretaceous, North Africa was transgressed many times by a southward extending arm of the Tethys Sea. These events produced a diverse set of coastal lithosomes preserved in part in the Cenomanian (99.0 to 93.5 Ma) Bahariya Formation. These deposits are well-exposed in the Bahariya Oasis (320 km southwest of Cairo) (Figure 5) and record a complex depositional history along a sheltered, low-energy, epeiric bight. The authors examined these deposits and the fossil biota contained within during two field seasons in 2000 and 2001. The Bahariya Bight coast developed along a very low-gradient coastal plain, producing a wide swath of paralic environments, consisting of lagoons, tidal flats, oyster reefs (Exogyra sp.), mangals, and tidal channels. The littoral zone was lined by a chain of low, sediment-starved, transgressive barrier islands. In areas of particularly low wave energy, the coast was directly colonized by mangrove vegetation. By the early Late Cretaceous, angiosperm mangrove and salt marsh species had not yet evolved. The mangrove flora along the Bahariya Bight was dominated by the halophytic tree-fern Weichselia reticulata. This plant was well-adapted to life along the coast. Woody prop roots helped stabilize it against waves and currents, and xeromorphic leaves reduced its need for fresh water (Alvin, 1974). Weichselia is known for its coastal affinity (Alvin, 1971; Barale, 1979; Daber, 1968; El Khayal, 1985; Lejal-Nicol and Dominik, 1990; Shinaq and Bandel, 1998; Smith et al., 2001) and Retallack and Dilcher (1981) believe that it formed a pantropical mangrove during the Early to early Late Cretaceous. Weichselia fossils are not known from post-Cenomanian deposits. A Late Cretaceous angiosperm invasion of the mangrove biome may have led to the fern’s extinction (Retallack and Dilcher, 1981). 1777 Lacovara et al., 2003 Figure 5. The Cenomanian Bahariya Formation is exposed in the Bahariya Oasis, located approximately 320 kilometers southwest of Cairo in the Western Desert of Egypt. (NASA satellite image) The Bahariya Bight coast appears to have been tide-dominated. Mud drapes indicate the likelihood of mesotidal conditions, whereas, fine-grained foreshore deposits point to a very lowenergy wave regime. Paleobarrier sands from the Bahariya Formation are super-mature (≈ 100% quartz) and show the isolation of this system from source material on the Eastern Saharan Craton. The inferred paucity of sediment in the paleolittoral system is corroborated by an abundance of glauconite in shoreface deposits, which is typically taken to indicate a low rate of clastic influx (see Harris and Whiting, 2000). The North African Tethys coast was situated near the paleoequator and Bahariya Formation deposits show evidence of an extremely productive coastal ecosystem. In addition to abundant plant remains, these deposits have yielded the remains of at least five dinosaurian genera (Smith et al., 2001; Stromer, 1915, 1931, 1932), 11 other tetrapod species (Stromer, 1914, 1925, 1933, 1934a, b, 1935, 1936), a rich ichthyofauna, including chondrichthyan and osteichthyan taxa (Peyer, 1925; Schaal, 1984; Stromer, 1925, 1927, 1936; Weiler, 1935; Werner, 1989), and an invertebrate assemblage that includes a recently described crab (Schweitzer et al., in press). MODERN/ANCEINT ANALOG Inlets dominate the Ten Thousand Islands coast. Barriers are small, frequently dissected, and often form only pocket beaches between mangrove promontories (Figure 6). Backbarrier mangrove islands are numerous and separated by a great number of tidal channels. Lightly vegetated tidal flats 1778 Lacovara et al., 2003 flank many of the islands and open-water lagoons exist towards the mainland. Overall, the landscape is a patchwork of littoral and paralic subenvironments. Figure 6. A portion of the Ten Thousand Islands coast. (NOS, NOAA air photo) The discontinuous coastal lithosomes of the Bahariya Formation reflect deposition under analogous conditions. In most cases it is not possible to follow a single facies for more than a hundred meters. Foreshore sands are thin and generally do not extend beyond an individual exposure. Tidal flat facies are common and often grade laterally into channel deposits. Open-water lagoon deposits are present, but typically thin (< 1m). Tidal channel deposits and mangrove paleosols are the two most common facies in the Bahariya Formation. They are typically about 10 to 100 cm thick and often alternate vertically. The resulting sequence records a two part cycle in which: 1) mangrove islands are dissected by tidal channels, and; 2) tidal channels are recolonized by mangroves (Figure 7). Foreshore deposits in the Bahariya Formation are generally found above paralic sediments, forming the classic transgressive sequence for barrier islands (sensu Kraft, 1971). In places, however, paralic deposits, containing plant fossils, overlie glauconitic shoreface sands (Figure 8). Following the Kraft Model, this sequence would represent discontinuous deposition and the boundary between the units would be interpreted as an unconformity. Close inspection of the outcrops, however, does not reveal an unconformity; the transition between these facies is 1779 Lacovara et al., 2003 gradational. This unusual juxtaposition of paleoenvironments can be explained by Parkinson’s (1989) model for the Ten Thousand Islands coast. The Cretaceous corollary is as follows: In the sheltered waters of the Cenomanian Bahariya Bight, wave-resistant tree-ferns (Weichselia reticulata) colonized the low-energy littoral zone. As sediment accumulated around the prop roots of Weichselia, mangrove islands prograded into the sea. Small portions of the coast regressed, even as sea level rose. Along reaches where wave-energy was slightly elevated, small barrier islands developed and migrated landward over paralic deposits. Abundant plant life along this warm, quiescent coast provided a broad trophic base for an extremely productive ecosystem. Figure 7. In the foreground, about 2 meters of exposure of the Bahariya Formation, showing tidal channel deposits above a mangrove paleosol. (Photo: K. Lacovara) Figure 8. The Bahariya Formation showing mangrove paleosol and tidal deposits over glauconitic shoreface sands. The paleosol contains woody plant remains, some of which extend through the contact, into the shoreface unit. (Photo: K. Lacovara) 1780 Lacovara et al., 2003 CONCLUSIONS In both the Holocene sediments of the Ten Thousand Islands and the Cretaceous sediments of the Bahariya Formation, two modes of coastal evolution in response to rising seas are recorded: transgressive and regressive. It is important to note that: 1) both modes occur simultaneously along different reaches of the same coast, and; 2) the resulting transgressive and regressive sequences do not represent a change in sea level trend, but rather represent local heterogeneities in wave climate. Although the evolution of the Ten Thousand Islands coast can be viewed as anomalous today, similar morphodynamic settings must have been widespread along the low-energy, lushly vegetated epeiric coasts of the Cretaceous. Without adequate modern analogs, the complex depositional history of these epeiric coasts might be undecipherable. Conversely, as we attempt to understand the ramifications of current global warming and sea level rise, Cretaceous coastal deposits provide an ideal representation of coastal response to end-member sea level and climatic conditions. ACKNOWLEDGEMENTS We would like to thank the Egyptian Geological Survey and Mining Authority and the government of Egypt for their support and cooperation. This research was funded in part by Cosmos Studios and MPH Entertainment. Thanks to Christopher R. Stotese and the PALEOMAP Project (www.scotese.com) for the use of their paleogeographic reconstructions. More information about this project is available at www.egyptdinos.org. REFERENCES Alvin, K.L., 1971, Weichselia reticulata (Stokes et Webb) Fontaine from the Wealden of Belgium, Memoires, Volume 166: Brussels, Institut Royal des Sciences Naturelles de Belgique, p. 33. —, 1974, Leaf anatomy of Weichselia based on fusainized material: Palaeontology, v. 17, p. 587-598. Barale, G., 1979, Decouverte de Weichselia reticulata (Stokes & Webb) Fontaine emend. Alvin, filicinee leptosporangiee dans le Cretace inferieur de la province de Lerida (Espange); implications stratigraphiques et paleoecologiques: Geobios, v. 12, p. 313-318. Bernard, H.A., and LeBlanc, R.J., Sr., 1965, Resume of the Quaternary geology of the northwestern Gulf of Mexico Provice, in Wright, H.E., and Frey, D.G., eds., The Quaternary of the United States: Princeton, N.J., Princeton University Press, p. 137-185. Daber, R., 1968, A Weichselia-Stiehleria-Matoniaceae community within the Quedlinburg estuary of lower Cretaceous age: Journal of the Linnean Society of London, Botany, v. 61, p. 75-85. Davis, R.A., Hine, A.C., and Shinn, E.A., 1992, Holocene coastal development on the Florida Peninsula, in Wehmiller, J.F., and Fletcher III, C.H., eds., Quaternary Coasts of the 1781 Lacovara et al., 2003 United States: Marine and Lacustrine Systems, Volume 48: Tulsa, Oklahoma, SEPM (Society for Sedimentary Geology), p. 193-212. El Khayal, A.A., 1985, Occurrence of characteristic Wealden fern (Weichselia reticulata) in the Wasia Formation, central Saudi Arabia.: Scripta Geologica, v. 79, p. 75-88. Enos, P., and Perkins, R.D., 1979, Evolution of Florida Bay from island stratigraphy: Geological Society of America Bulletin, v. 90, p. 59-83. Haq, B.U., Hadenbol, J., and Vail, P.R., 1987, The chronology of fluctuating sea level since the Triassic: Science, v. 235, p. 1156-1167. Harris, L.C., and Whiting, B.M., 2000, Sequence-stratigraphic significance of Miocene to Pliocene glauconite-rich layers, on- and offshore of the US Mid-Atlantic margin: Sedimentary Geology, v. 134, p. 129-147. Kraft, J.C., 1971, Sedimentary environment, facies pattern, and geologic history of a Holocene marine transgression: Geological Society America Bulletin, v. 82, p. 2131-2158. Lejal-Nicol, A., and Dominik, W., 1990, Sur la paleoflore a Weichseliaceae et a angiospermes du Cenomanien de la region de Bahariya (Egypte du Sud-Ouest): Berliner Geowissenschaftliche Abhandlungen, Reihe A: Geologie und Palaeontologie, v. 120, p. 957-991. Parkinson, R.W., 1987, Holocene sedimentation and coastal response to rising sea level along a subtropical low energy coast, Ten Thousand Islands, Southwest Florida [Dissertation thesis]: Coral Gables, Florida, University of Miami. —, 1989, Decelerating Holocene sea-level rise and its influence on southwest Florida coastal evolution: a transgressive / regressive stratigraphy: Journal of Sedimentary Petrology, v. 59, p. 960-972. Peyer, B., 1925, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wüsten Agyptens. II. Wirbeltier-Reste der Baharije-Stufe (unterstes Cenoman). 6. Die Ceratodus-Funde.: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematischenaturwissenschaftliche Abteilung., v. 30, p. 1-23. Retallack, G., and Dilcher, D.L., 1981, A coastal hypothesis for the dispersal and rise to dominance of flowering plants, in Niklas, K.J., ed., Paleobotany, Paleoecology, and Evolution, Volume 2: New York, Praeger, p. 27-77. Schaal, S., 1984, Oberkretazische Osteichthyes (Knochenfische) aus dem Bereich von Bahariya und Kharga, Ägypten, und ihre Aussagen zur Palökologie und Stratigraphie: Berlin, D. Reimer, 79 , ix leaves of plates p. Scholl, D.W., Graighead, F.C., and Stuiver, M., 1969, Florida submergence curve revisited: its relation to sedimentation rates: Science, v. 163, p. 562-564. Schweitzer, C.E., Lacovara, K.J., Smith, J.B., Lamanna, M.C., Lyon, M.A., and Attia, Y., in press, Mangrove-dwelling crabs (Brachyura: Necrocarcinidae?) associated with dinosaurs from the Upper Cretaceous (Cenomanian) of Egypt: Journal of Paleontology, p. 23 p. ms. Scotese, C.R., 2001, Atlas of Earth History, Volume 1, Paleogeography: Arlington, Texas, PALEOMAP Project, 52 p. Shinaq, R., and Bandel, K., 1998, The flora of an estuarine channel margin in the Early Cretaceous of Jordan, Freiberger Forschungshefte C 474, Paläontologie, Stratigraphie, Fazies - Heft 6: Freiberger, p. 39-57. 1782 Lacovara et al., 2003 Smith, J.B., Lamanna, M.C., Lacovara, K.J., Dodson, P., Smith, J.R., Poole, J.C., Giegengack, R., and Attia, Y., 2001, A giant sauropod dinosaur from an Upper Cretaceous mangrove deposit in Egypt: Science, v. 292, p. 1704-1706. Stromer, E., 1914, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 1. Einleitung und 2. Libycosuchus: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathemetsch-physikalische Klasse, v. 27, p. 1-16. —, 1915, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 3. Das Original des Theropoden Spinosaurus aegyptiacus: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathemetsch-naturwissenschaftliche Abteilung, v. 28, p. 1-32. —, 1925, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 7. Stomatosuchus inermis Stromer, ein schwach bezanhter Krokodilier und 8. Ein Skelettrest des Pristiden Onchopristis numidus Haug sp.: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathemetsch-naturwissenschaftliche Abteilung, v. 30, p. 1-22. —, 1927, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 9. Die Plagiostomen, mit einem Anhang uber kano- und mesozoische Ruckenflossenstacheln von Elasmobranchiern: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathemetschnaturwissenschaftliche Abteilung, v. 31, p. 1-64. —, 1931, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 10. Ein Skelett-Rest van Carcharodontosaurus nov. gen.: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathemetsch-naturwissenschaftliche Abteilung, Neue Folge, v. 9, p. 123. —, 1932, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 11. Sauropoda: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathemetsch-naturwissenschaftliche Abteilung, Neue Folge, v. 10, p. 1-21. —, 1933, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 12. Die procolen Crocodilia: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathemetschnaturwissenschaftliche Abteilung, Neue Folge, v. 15, p. 1-55. —, 1934a, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 14. Testudinata: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathemetsch-naturwissenschaftliche Abteilung, Neue Folge, v. 25, p. 1-26. —, 1934b, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Ägyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 13. Dinosauria: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Abteilung, Neue Folge, v. 22, p. 1-79. —, 1935, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, II. Wirbeltier-Reste der Baharijestufe (unterstes Cenoman), 15. Plesiosauria: Abhandlungen 1783 Lacovara et al., 2003 der Bayerischen Akademie der Wissenschaften, Mathemetsch-naturwissenschaftliche Abteilung, Neue Folge, v. 26, p. 1-56. —, 1936, Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Ägyptens, VII. Baharîje-Kessel und -Stufe mit deren fauna und flora eine ergänzende zusammenfassung: Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematischnaturwissenschaftliche Abteilung, Neue Folge, v. 33, p. 3-102. Tanner, W.F., 1960, Florida coastal classification: Transport: Gulf Coast Association of Geologic Society, v. 10, p. 259-266. Thomlinson, B.P., 1986, The Botany of Mangroves: Cambridge, UK, Cambridge University Press, 419 p. Weiler, W., 1935, Ergebnisse der Forchungsreisen Prof. E. Stromers in den Wüsten Agyptens, II. Wirbeltierreste der Baharije-Stufe (unterstes Cenoman), 16. Neue Untersuchungen an den Fischesten. Abhandlungen der Bayerischen Akademie der Wissenschaften: Mathematischenaturwissenschaftliche Abteilung Neue Folge Hefte, v. 32, p. 1-57. Werner, C., 1989, Die Elasmobranchier-Fauna des Gebel Dist Member der Bahariya Formation (Obercenoman) der Oase Bahariya, Agypten: Palaeo Ichthyologica, v. 5, p. 1-112. Key words: Egypt, Florida, Bahariya Formation, Ten Thousand Islands, mangrove, Weichselia, Cretaceous, Cenomanian, Tethys, epeiric sea 1784