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Relationship between climate and plant functional types in the Congo Basin Rita Wania, Charly Favier and the CoForChange Consortium [email protected] L’Institut des Sciences de l’Evolution Montpellier, France 1 / 34 The Congo Basin 2 / 34 Plant Functional Types (PFTs) 3 / 34 Leaf shape: broad-leaved vs needle-leaved 4 / 34 Phenology: evergreen vs deciduous 5 / 34 Growth form: tree vs shrub 6 / 34 Climate: boreal vs tropical 7 / 34 Shade tolerance: tolerant vs intolerant 8 / 34 Current PFTs in LPJ-GUESS 1. Boreal shade-tolerant needleleaved evergreen tree 2. Boreal shade-intolerant needleleaved evergreen tree 3. Boreal shade-intolerant needleleaved summergreen tree 4. Temperate shade-tolerant broadleaved summergreen tree 5. Temperate shade-intolerant broadleaved summergreen tree 6. Temperate shade-intolerant broadleaved evergreen tree 7. Tropical shade-tolerant broadleaved evergreen tree 8. Tropical shade-intolerant broadleaved evergreen tree 9. Tropical shade-intolerant broadleaved raingreen tree 10. C3 Grasses 11. C4 Grasses 9 / 34 LPJ-GUESS 10 / 34 Which PFTs do the observations support? 11 / 34 Classification Guilds I P = Pioneer I NPLD = Non-Pioneer Light Demander I SB = Shade Bearer Phenology I evergreen I deciduous 12 / 34 Classification Guilds Phenology I P = Pioneer I NPLD = Non-Pioneer Light Demander I SB = Shade Bearer I evergreen I deciduous PFTs 1. Evergreen P 2. Evergreen NPLD 3. Evergreen SB 4. Deciduous P 5. Deciduous NPLD 6. Deciduous SB 12 / 34 Available data from forest inventories Inventory data Coordinates 13.3E, 13.3E, 13.3E, 13.3E, 13.3E, ... 4.1N 4.1N 4.1N 4.1N 4.1N Size (ha) 0.5 0.5 0.5 0.5 0.5 Diameter (cm) 40 60 60 80 100 No Guild Phenology 2 1 1 1 1 NPLD NPLD NPLD SB SB deciduous deciduous deciduous non-class. evergreen Wood density (g cm−3 ) 0.49 0.49 0.51 NA 0.64 13 / 34 Available data from forest inventories Inventory data Coordinates 13.3E, 13.3E, 13.3E, 13.3E, 13.3E, ... I I I I 4.1N 4.1N 4.1N 4.1N 4.1N Size (ha) 0.5 0.5 0.5 0.5 0.5 Diameter (cm) 40 60 60 80 100 No Guild Phenology 2 1 1 1 1 NPLD NPLD NPLD SB SB deciduous deciduous deciduous non-class. evergreen Wood density (g cm−3 ) 0.49 0.49 0.51 NA 0.64 Region: 13–19◦ E, 0–5◦ N 0.5-ha plots → 67,635 Plots with >21 trees → 52,166 Plots with trees with wood density estimation → 28,683 13 / 34 Can the variation of vegetation in the CoForChange study area be explained by climate? 14 / 34 Can the variation of vegetation in the CoForChange study area be explained by climate? Previous research attributed the distribution of species and functional traits to geology rather than climate! (Fayolle et al., subm.) 14 / 34 Climate clusters 15 / 34 Climate data used – WorldClim air temperature Monthly air temperature 1950–2000 @ 10kmx10km 16 / 34 Climate data used – FAO LocClim precipitation Monthly precipitation 1961–1990 @ 10kmx10km 17 / 34 Climate data used – CRU % cloud cover Monthly percentage cloud cover 1961–1990 @ 0.5◦ x0.5◦ 18 / 34 Clustering setup I Use R-function kmeans I Based only on climate data I Use Pr, Temp, Cld I Use 1, 2 or all 3 variables at a time I Use JJA, JJASO, DJF, annual 19 / 34 Spatial climate clusters Based on T (DJF), P (DJF) and cloud (annual). 20 / 34 What constitutes a good cluster? 21 / 34 What constitutes a good cluster? 21 / 34 What constitutes a good cluster? 20 10 0 Frequency 30 40 Histogram 40 50 60 70 80 90 100 % Cloud cover 21 / 34 What constitutes a good cluster? 21 / 34 What constitutes a good cluster? 21 / 34 What constitutes a good cluster? 21 / 34 Statistical measure: Coefficient of Variance (CV) CVcluster = X m=months σm µm (1) 22 / 34 What constitutes a good cluster? 80 80 70 70 40 40 50 50 60 60 % cloud cover % cloud cover 80 80 70 70 60 60 50 50 40 40 22 44 66 88 1010 1212 22 Months Months 44 66 88 1010 1212 Months Months CVcluster =440.062 Cluster Cluster 90 90 CV cluster33= 0.071 Cluster Cluster 90 90 % cloud cover % cloud cover 90 90 Cluster Cluster22 90 90 Cluster Cluster11 23 / 34 Spatial climate clusters Based on T (DJF), P (DJF) and cloud (annual). 24 / 34 Climate in each cluster: Cloud cover 25 / 34 Climate in each cluster: Precipitation 26 / 34 Climate clusters applied to inventory data 27 / 34 Apply cluster groups to vegetation variables I % deciduous/evergreen I % pioneer, % NPLD, % SB I % deciduous pioneer, % deciduous NPLD, % deciduous SB I % evergreen pioneer, % evergreen NPLD, % evergreen SB 28 / 34 Percentage Basal area: Deciduous PFTs Basal area, % decid PFTs 100 cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● 20 40 60 80 ● ● ● ● ● ● 0 ● ● ● ● ● ● 1 2 3 4 5 6 29 / 34 Percentage Basal area: Deciduous PFTs Basal area, % decid PFTs 100 cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● 20 40 60 80 ● ● ● ● ● ● 0 ● ● ● ● ● ● 1 2 3 4 5 6 29 / 34 Percentage Basal area: Deciduous PFTs Basal area, % decid PFTs 100 cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● 20 40 60 80 ● ● ● ● ● ● 0 ● ● ● ● ● ● 1 2 3 4 5 6 29 / 34 Percentage Basal area: Deciduous PFTs Basal area, % decid PFTs 100 cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● 20 40 60 80 ● ● ● ● ● ● 0 ● ● ● ● ● ● 1 2 3 4 5 6 29 / 34 Shade bearers Basal area, % decid Shade tolerant ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 2 3 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 0 0 10 20 20 40 30 40 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 cldAnn−tasDJF−prDJF−n6−best−pr ● 50 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● 80 100 cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 70 Basal area, % ever Shade tolerant ● ● ● ● ● ● ● ● 1 2 3 4 5 6 1 4 5 6 30 / 34 Shade bearers Basal area, % decid Shade tolerant ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 2 3 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 0 0 10 20 20 40 30 40 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 cldAnn−tasDJF−prDJF−n6−best−pr ● 50 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● 80 100 cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 70 Basal area, % ever Shade tolerant ● ● ● ● ● ● ● ● 1 2 3 4 5 6 1 4 5 6 30 / 34 Shade bearers Basal area, % decid Shade tolerant ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 2 3 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 0 0 10 20 20 40 30 40 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 cldAnn−tasDJF−prDJF−n6−best−pr ● 50 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● 80 100 cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 70 Basal area, % ever Shade tolerant ● ● ● ● ● ● ● ● 1 2 3 4 5 6 1 4 5 6 30 / 34 Shade bearers Basal area, % decid Shade tolerant ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 2 3 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 0 0 10 20 20 40 30 40 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 cldAnn−tasDJF−prDJF−n6−best−pr ● 50 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● 80 100 cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 70 Basal area, % ever Shade tolerant ● ● ● ● ● ● ● ● 1 2 3 4 5 6 1 4 5 6 30 / 34 Basal area, % decid Pioneer cldAnn−tasDJF−prDJF−n6−best−pr 100 Basal area, % ever Pioneer cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● ● ● ● ● ● ● 1 2 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 80 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 40 ● ● ● ● ● ● ● 0 20 ● ● ● ● ● ● ● ● ● ● ● 0 20 40 60 80 100 Pioneers 3 4 5 6 1 2 3 4 5 6 31 / 34 Basal area, % decid Pioneer cldAnn−tasDJF−prDJF−n6−best−pr 100 Basal area, % ever Pioneer cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● ● ● ● ● ● ● 1 2 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 80 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 40 ● ● ● ● ● ● ● 0 20 ● ● ● ● ● ● ● ● ● ● ● 0 20 40 60 80 100 Pioneers 3 4 5 6 1 2 3 4 5 6 31 / 34 Basal area, % decid Pioneer cldAnn−tasDJF−prDJF−n6−best−pr 100 Basal area, % ever Pioneer cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● ● ● ● ● ● ● 1 2 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 80 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 40 ● ● ● ● ● ● ● 0 20 ● ● ● ● ● ● ● ● ● ● ● 0 20 40 60 80 100 Pioneers 3 4 5 6 1 2 3 4 5 6 31 / 34 Basal area, % decid Pioneer cldAnn−tasDJF−prDJF−n6−best−pr 100 Basal area, % ever Pioneer cldAnn−tasDJF−prDJF−n6−best−pr ● ● ● ● ● ● ● ● ● ● 1 2 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 80 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 40 ● ● ● ● ● ● ● 0 20 ● ● ● ● ● ● ● ● ● ● ● 0 20 40 60 80 100 Pioneers 3 4 5 6 1 2 3 4 5 6 31 / 34 Non-pioneer light demanding (NPLD) Basal area, % ever NPLD Basal area, % decid NPLD cldAnn−tasDJF−prDJF−n6−best−pr cldAnn−tasDJF−prDJF−n6−best−pr ● 80 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 40 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 0 0 20 20 40 60 ● ● ● ● ● ● 1 2 3 4 5 6 1 2 3 4 5 6 32 / 34 Non-pioneer light demanding (NPLD) Basal area, % ever NPLD Basal area, % decid NPLD cldAnn−tasDJF−prDJF−n6−best−pr cldAnn−tasDJF−prDJF−n6−best−pr ● 80 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 40 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 0 0 20 20 40 60 ● ● ● ● ● ● 1 2 3 4 5 6 1 2 3 4 5 6 32 / 34 Non-pioneer light demanding (NPLD) Basal area, % ever NPLD Basal area, % decid NPLD cldAnn−tasDJF−prDJF−n6−best−pr cldAnn−tasDJF−prDJF−n6−best−pr ● 80 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 40 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 0 0 20 20 40 60 ● ● ● ● ● ● 1 2 3 4 5 6 1 2 3 4 5 6 32 / 34 Non-pioneer light demanding (NPLD) Basal area, % ever NPLD Basal area, % decid NPLD cldAnn−tasDJF−prDJF−n6−best−pr cldAnn−tasDJF−prDJF−n6−best−pr ● 80 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 60 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 40 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● 0 0 20 20 40 60 ● ● ● ● ● ● 1 2 3 4 5 6 1 2 3 4 5 6 32 / 34 Summary I Deciduous PFTs contribute between 35 and 70% to basal area. 33 / 34 Summary I I Deciduous PFTs contribute between 35 and 70% to basal area. High deciduousness was related to lower T, stronger JJA dry season and higher cloud cover in that dry season. 33 / 34 Summary I I I Deciduous PFTs contribute between 35 and 70% to basal area. High deciduousness was related to lower T, stronger JJA dry season and higher cloud cover in that dry season. Evergreen shade bearers showed the opposite pattern to decidousness. 33 / 34 Summary I I I I Deciduous PFTs contribute between 35 and 70% to basal area. High deciduousness was related to lower T, stronger JJA dry season and higher cloud cover in that dry season. Evergreen shade bearers showed the opposite pattern to decidousness. Deciduous pioneers were higher in cooler regions, with a stronger JJA dry season and an overall higher cloud cover. 33 / 34 Summary I I I I I Deciduous PFTs contribute between 35 and 70% to basal area. High deciduousness was related to lower T, stronger JJA dry season and higher cloud cover in that dry season. Evergreen shade bearers showed the opposite pattern to decidousness. Deciduous pioneers were higher in cooler regions, with a stronger JJA dry season and an overall higher cloud cover. Decidous NPLDs were more abundant in warmer regions with a milder JJA dry season, but a stronger DJF season and a low cloud cover. 33 / 34 Summary I I I I I I Deciduous PFTs contribute between 35 and 70% to basal area. High deciduousness was related to lower T, stronger JJA dry season and higher cloud cover in that dry season. Evergreen shade bearers showed the opposite pattern to decidousness. Deciduous pioneers were higher in cooler regions, with a stronger JJA dry season and an overall higher cloud cover. Decidous NPLDs were more abundant in warmer regions with a milder JJA dry season, but a stronger DJF season and a low cloud cover. Pioneers and NPLD are both light demanding, but they occur in different climate zones. Pioneers don’t mind the higher cloud cover, probably because they depend more on forest openings and the sunlight reaching the forest floor, whereas NPLDs seem to do better in low cloud cover regions as they are light-demanders as mature trees. 33 / 34 Summary I I I I I I Deciduous PFTs contribute between 35 and 70% to basal area. High deciduousness was related to lower T, stronger JJA dry season and higher cloud cover in that dry season. Evergreen shade bearers showed the opposite pattern to decidousness. Deciduous pioneers were higher in cooler regions, with a stronger JJA dry season and an overall higher cloud cover. Decidous NPLDs were more abundant in warmer regions with a milder JJA dry season, but a stronger DJF season and a low cloud cover. Pioneers and NPLD are both light demanding, but they occur in different climate zones. Pioneers don’t mind the higher cloud cover, probably because they depend more on forest openings and the sunlight reaching the forest floor, whereas NPLDs seem to do better in low cloud cover regions as they are light-demanders as mature trees. 33 / 34 Conclusion Overall, we think that the vegetation distribution in the CoForChange study area can be (at least partially) explained by climatic variables. 34 / 34 Conclusion Overall, we think that the vegetation distribution in the CoForChange study area can be (at least partially) explained by climatic variables. However, first runs with LPJ-GUESS do not show a climatic effect on simulated vegetation composition between the six clusters. 34 / 34 Conclusion Overall, we think that the vegetation distribution in the CoForChange study area can be (at least partially) explained by climatic variables. However, first runs with LPJ-GUESS do not show a climatic effect on simulated vegetation composition between the six clusters. The question is, is the climatic effect we see for real or do the climate clusters coincide with disturbance history that cause the different vegetation pattern? 34 / 34
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